Detectives Nick O'Malley and Kate Benson work in Special Unit 2, a secret precinct of the Chicago Police Department whose sole charge is to protect the city's citizens from Links, a malicious paranormal species that is the missing link between man and beast.

When the local FBI office receives a letter from a terrorist known only as 'The Citizen', it's quickly determined that he's planning his next act at the Miss America beauty pageant. Because tough-as-nails Gracie Hart is the only female Agent at the office, she's chosen to go undercover as the contestant from New Jersey.